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TP103 – Simplified metadata

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Dataset

Database Code
TP103
Title
Species interactions during succession in the western Cascade Range of Oregon, 1990 to present
Abstract
The factors that contribute to plant species establishment and decline following disturbance determine the rates and patterns of successional change of a system. In this long-term field experiment, we test the commonly held assumption that competition for space or resources by dominant species determines the outcome of succession. Specifically, we examine the population- and community-level consequences of removing one or more potentially dominant species from the post-disturbance community after clearcut logging and burning of a mature/old-growth Douglas-fir forest. Experimental treatments include: (1) removal of early-seral annual, Senecio sylvaticus, or perennial, Epilobium angustifolium—or both—to test the influences of these early-seral dominants on subsequent community development; (2) removal of all species except Senecio or Epilobium, to test whether the decline of these early-seral dominants is driven by competitive displacement; or (3) removal of shade-tolerant forest species that dominate subsequent stages of succession—Rubus ursinus or Berberis nervosa plus Gaultheria shallon—to test the influences of these long-lived perennials on understory development. The experiment is a randomized block design comprising eight removal treatments plus a control replicated in each of 25 blocks. Removal (reduction in competition) is achieved by removing seedlings or vegetative stems annually from a treatment area of 2.5 x 2.5 m. Sample plots (1 x 1 m) centered within these are used to estimate cover of all vascular plant species and, for the first 8 yr of the experiment, stem density and height, facilitating estimates of above-ground biomass. Pre-harvest data were collected in 1990, logging/burning occurred in 1991, and removal treatments and post-treatment sampling were initiated in 1992. Six of the nine experimental treatments were terminated between 1996 and 1998, with loss of early-seral Senecio and Epilobium from the system. The remaining three treatments (removal of Rubus, removal of Berberis plus Gaultheria, and the control) have been sampled annually since 1992. All regeneration of trees within the experimental area has occurred naturally. In September 2018, the study site was subjected to a controlled burn to limit spread of the Terwilliger Fire from the north. In 2019, plots were assessed for burn severity. Species' removals were terminated, but cover measurements continue in the plots.
Keywords
community dynamics; trophic structure; succession; primary production; species interactions; competition; disturbance; burning; timber harvest; clearcutting; Long-Term Ecological Research (LTER); primary production; disturbance
Date begin
1990-07-01
Date end
2024-07-03
Principal Investigator
Charles B. Halpern

Entities

  1. Species Cover, Density, and Biomass Measurement Data
  2. Species Removals (counts)
  3. Species Cover, Density, and Biomass Measurements Original Fieldsheet Format
  4. Species Cover, Density, and Biomass Measurements Modified Fieldsheet Format
  5. Equations used for calculating biomass
  6. Estimated biomass using measurement data and equations
  7. Assessment of 2018 burn severity

1. Species Cover, Density, and Biomass Measurement Data - Attribute list

Species Cover, Density, and Biomass Measurement Data - attributes
Attribute Type Domain Minimum Maximum Unit
DBCODE varchar enum
ENTITY numeric(1.000000) range 1.0000 1.0000 number
STUDY varchar enum
YEAR numeric(1.000000) range 1990.0000 2024.0000 number
SDATE datetime freetext
SEASON varchar enum
BLOCK numeric(1.000000) range 1.0000 25.0000 number
PLOT numeric(1.000000) range 1.0000 9.0000 number
TMT varchar enum
SPECIES varchar enum
STRATUM varchar enum
COV1_4TH numeric(0.100000) range 0.0000 100.0000 percent
COVER numeric(0.100000) range 0.0000 100.0000 percent
DENSITYS numeric(1.000000) range 0.0000 3300.0000 number
DENSITYM numeric(1.000000) range 0.0000 105.0000 number
DBA numeric(0.100000) range 0.0000 100.0000 centimeters
HT numeric(0.100000) range 0.0000 145.0000 centimeters
HTV numeric(0.100000) range 0.0000 330.0000 centimeters
REC_NO numeric(1.000000) range 1.0000 1135.0000 number

2. Species Removals (counts) - Attribute list

Species Removals (counts) - attributes
Attribute Type Domain Minimum Maximum Unit
DBCODE varchar enum
ENTITY numeric(1.000000) range 2.0000 2.0000 number
STUDY varchar enum
SDATE datetime freetext
RDATE datetime freetext
SEASON varchar enum
BLOCK numeric(1.000000) range 1.0000 25.0000 number
PLOT numeric(1.000000) range 1.0000 9.0000 number
TMT varchar enum
SPECIES varchar enum
NREMBUF numeric(1.000000) range 0.0000 1354.0000 number
NREMPLT numeric(1.000000) range 0.0000 486.0000 number
NOTES varchar freetext

3. Species Cover, Density, and Biomass Measurements Original Fieldsheet Format - Attribute list

Species Cover, Density, and Biomass Measurements Original Fieldsheet Format - attributes
Attribute Type Domain Minimum Maximum Unit
DBCODE varchar enum
ENTITY numeric(1.000000) range 3.0000 3.0000 number
STUDY varchar enum
YEAR numeric(1.000000) range 1990.0000 1996.0000 number
SDATE datetime freetext
SEASON varchar enum
BLOCK numeric(1.000000) range 1.0000 25.0000 number
PLOT numeric(1.000000) range 1.0000 9.0000 number
TMT varchar enum
SPECIES varchar enum
STRATUM varchar enum
LC numeric(1.000000) range 1.0000 165.0000 number
COV1_4TH numeric(0.100000) range 0.0000 100.0000 percent
COVER numeric(0.100000) range 0.0000 100.0000 percent
DENSITYS numeric(1.000000) range 0.0000 3300.0000 number
DENSITYM numeric(1.000000) range 0.0000 120.0000 number
STEM1BV1 numeric(0.100000) range 0.0000 130.0000 number
STEM1BV2 numeric(0.100000) range 0.0000 130.0000 number
STEM2BV1 numeric(0.100000) range 0.0000 330.0000 number
STEM2BV2 numeric(0.100000) range 0.0000 150.0000 number
STEM3BV1 numeric(0.100000) range 0.0000 130.0000 number
STEM3BV2 numeric(0.100000) range 0.0000 130.0000 number
STEM4BV1 numeric(0.100000) range 0.0000 130.0000 number
STEM4BV2 numeric(0.100000) range 0.0000 130.0000 number
STEM5BV1 numeric(0.100000) range 0.0000 130.0000 number
STEM5BV2 numeric(0.100000) range 0.0000 130.0000 number

4. Species Cover, Density, and Biomass Measurements Modified Fieldsheet Format - Attribute list

Species Cover, Density, and Biomass Measurements Modified Fieldsheet Format - attributes
Attribute Type Domain Minimum Maximum Unit
DBCODE varchar enum
ENTITY numeric(1.000000) range 4.0000 4.0000 number
STUDY varchar enum
YEAR numeric(1.000000) range 1997.0000 2002.0000 number
SDATE datetime freetext
SEASON varchar enum
BLOCK numeric(1.000000) range 1.0000 25.0000 number
PLOT numeric(1.000000) range 1.0000 9.0000 number
TMT varchar enum
SPECIES varchar enum
LC numeric(1.000000) range 1.0000 10.0000 number
COV1_4TH numeric(0.100000) range 0.0000 100.0000 percent
COVER numeric(0.100000) range 0.0000 100.0000 percent
DENSITYS numeric(1.000000) range 0.0000 2000.0000 number
DENSITYM numeric(1.000000) range 0.0000 70.0000 number
STEM1BV1 numeric(0.100000) range 0.1000 150.0000 number
STEM1BV2 numeric(0.100000) range 0.1000 150.0000 number
STEM1NO numeric(1.000000) range 1.0000 1000.0000 number
STEM2BV1 numeric(0.100000) range 0.1000 150.0000 number
STEM2BV2 numeric(0.100000) range 0.1000 150.0000 number
STEM2NO numeric(1.000000) range 1.0000 1000.0000 number
STEM3BV1 numeric(0.100000) range 0.1000 150.0000 number
STEM3BV2 numeric(0.100000) range 0.1000 150.0000 number
STEM3NO numeric(1.000000) range 1.0000 1000.0000 number

5. Equations used for calculating biomass - Attribute list

Equations used for calculating biomass - attributes
Attribute Type Domain Minimum Maximum Unit
DBCODE varchar enum
ENTITY numeric(1.000000) range 5.0000 5.0000 number
STUDY varchar enum
YEAR numeric(1.000000) range 1992.0000 2024.0000 number
SPECIES varchar enum
EQUATION varchar freetext
ALTEQU varchar enum

6. Estimated biomass using measurement data and equations - Attribute list

Estimated biomass using measurement data and equations - attributes
Attribute Type Domain Minimum Maximum Unit
DBCODE varchar enum
ENTITY numeric(1.000000) range 6.0000 6.0000 number
STUDY varchar enum
YEAR numeric(1.000000) range 1992.0000 2024.0000 number
SEASON varchar enum
BLOCK numeric(1.000000) range 1.0000 25.0000 number
PLOT numeric(1.000000) range 1.0000 9.0000 number
TMT varchar enum
SPECIES varchar enum
B_VAL numeric(1) range 1.0000 8.0000 number
BIOMASS numeric(0.010000) range 0.0000 2279755.7500 grams per square meter

7. Assessment of 2018 burn severity - Attribute list

Assessment of 2018 burn severity - attributes
Attribute Type Domain Minimum Maximum Unit
DBCODE varchar enum
ENTITY numeric(1.000000) range 7.0000 7.0000 number
STUDY varchar enum
YEAR numeric(1.000000) range 2019.0000 2019.0000 number
SDATE datetime freetext
BLOCK numeric(1.000000) range 1.0000 25.0000 number
PLOT numeric(1.000000) range 1.0000 9.0000 number
TMT varchar enum
TREES_LIVE numeric(1) range 0.0000 12.0000 number
TREES_DEAD numeric(1) range 0.0000 11.0000 number
PROP_DEAD numeric(1) range 0.0000 100.0000 number
PCT_BURN numeric(1) range 0.0000 100.0000 percent
PCT_BURN_ADJ numeric(1) range 0.0000 100.0000 percent
BURN_CLASS varchar enum
BURN_COMMENT varchar freetext

Attribute definitions

ALTEQU
Alternative equation within a species for calculating biomass
BIOMASS
Estimated biomass
BLOCK
Sample block (1-25)
BURN_CLASS
Burn class
BURN_COMMENT
Description of burn conditions
B_VAL
Unique biomass value for a given year, season, block, plot, tmt, species.
COV1_4TH
Cover in the "quarter" plot (0.25 x 0.25 m), used to subsample when species density was high
COVER
Cover in the 1 m2 plot
DBA
Basal diameter
DBCODE
Database code
DENSITYM
Density of mature stems/sprouts (1 m2 plot)
DENSITYS
Density of seedlings (1 m2 plot)
ENTITY
Entity number
EQUATION
Explicit equation used for calculating biomass. mht = height, mcov= cover, mdba = bottom diameter
HT
Total height (including inflorescence if present)
HTV
Height of vegetative portion
LC
Line count
NOTES
Comments
NREMBUF
Number of stems removed from 75 cm buffer
NREMPLT
Number of stems removed from 1 m2 plot
PCT_BURN
Percentage of ground surface burned or charred within the plot
PCT_BURN_ADJ
Percentage of ground surface burned or charred within 0.5 m of the plot edge
PLOT
Sample plot w/in block (1-9)
PROP_DEAD
Proportion of trees within 2.5 m of the plot that are dead (undefined if there are no trees)
RDATE
Date of species removal (yyyymmdd)
REC_NO
Unique record number for a given year, season, block, plot, tmt, species and stratum
SDATE
Vegetation sampling date
SEASON
Sampling season (1 - 3)
SPECIES
Species acronym according to Garrison et al. 1974
STEM1BV1
Biomass parameter 1 of stem1
STEM1BV2
Biomass parameter 2 of stem1
STEM1NO
Number of stems containing the corresponding bv1 and bv2 values
STEM2BV1
Biomass parameter 1 of stem2
STEM2BV2
Biomass parameter 2 of stem2
STEM2NO
Number of stems containing the corresponding bv1 and bv2 values
STEM3BV1
Biomass parameter 1 of stem3
STEM3BV2
Biomass parameter 2 of stem3
STEM3NO
Number of stems containing the corresponding bv1 and bv2 values
STEM4BV1
Biomass parameter 1 of stem4
STEM4BV2
Biomass parameter 2 of stem4
STEM5BV1
Biomass parameter 1 of stem5
STEM5BV2
Biomass parameter 2 of stem5
STRATUM
Pre-harvest tree stratum: (O)verstory, (S)ubcanopy, (U)nderstory
STUDY
Study site location
TMT
Treatment code (A-I)
TREES_DEAD
Number of dead trees within 2.5 m of the plot
TREES_LIVE
Number of live trees within 2.5 m of the plot
YEAR
Sampling year

Enumerated domains

Enumerated domain for attribute: DBCODE

Codes for DBCODE
Code Description
TP103 FSDB database code TP103

Enumerated domain for attribute: STUDY

Codes for STUDY
Code Description
Starrbright Starrbright Timber Sale

Enumerated domain for attribute: SEASON

Codes for SEASON
Code Description
U Unknown season of sampling
1 First season of sampling per year (usually june)
2 Second season of sampling per year (usually july)
3 Third season of sampling per year (usually august)

Enumerated domain for attribute: TMT

Codes for TMT
Code Description
A Control
B Senecio sylvaticus removed
C Epilobium angustifolium removed
D Senecio sylvaticus and Epilobium angustifolium removed
E Senecio sylvaticus and community removed
F Epilobium angustifolium and community removed
G Community removed
H Rubus ursinus removed
I Berberis nervosa and Gaultheria shallon removed

Enumerated domain for attribute: SPECIES

Codes for SPECIES
Code Description
BORAGI Boraginaceae
LILIAC Liliaceae
ABGR Abies grandis
ACCI Acer circinatum
ACMA Acer macrophyllum
ACTR Achlys triphylla
ADBI Adenocaulon bicolor
AGGR Agoseris grandiflora
AGHE Agoseris heterophylla
AGROS Agrostis
AGEX Agrostis exarata
AGTE Agrostis tenuis
AICA Aira caryophyllea
AMAL Amelanchier alnifolia
ANMA Anaphalis margaritacea
ANDE Anemone deltoidea
ANLY2 Anemone lyallii
ARME Arbutus menziesii
ARCO3 Arctostaphylos columbiana
AREL Arrhenatherum elatius
ASCA3 Asarum caudatum
BENE Berberis nervosa
BROMU Bromus
BRVU Bromus vulgaris
CASC2 Campanula scouleri
CARDA Cardamine
CAOL Cardamine oligosperma
CAAM Carex amplifolia
CEANO Ceanothus
CEIN Ceanothus integerrimus
CESA Ceanothus sanguineus
CEVE Ceanothus velutinus
CERAS Cerastium
CEVU Cerastium vulgatum
CHME Chimaphila menziesii
CHUM Chimaphila umbellata
CHLE2 Chrysanthemum leucanthemum
CIRSI Cirsium
CIAR Cirsium arvense
CIVU Cirsium vulgare
CLUN Clintonia uniflora
COHE Collomia heterophylla
COCA2 Conyza canadensis
CONU Cornus nuttallii
CRCA Crepis capillaris
DEEL Deschampsia elongata
ELGL Elymus glaucus
EPILO Epilobium
EPMI Epilobium minutum
EPPA Epilobium paniculatum
EPWA Epilobium watsonii
EQTE Equisetum telmateia
FESTU Festuca
FEMY Festuca myuros
FEOC Festuca occidentalis
FRVE Fragaria vesca
GALIU Galium
GATR Galium triflorum
GAFR Garrya fremontii
GASH Gaultheria shallon
GNPU Gnaphalium purpureum
GNUL Gnaphalium uliginosum
GOOB Goodyera oblongifolia
HIAL Hieracium albiflorum
HOLA Holcus lanatus
HODI Holodiscus discolor
HYPE Hypericum perforatum
HYRA Hypochaeris radicata
LAMU Lactuca muralis
LASE Lactuca serriola
LIBO2 Linnaea borealis
LISTE Listera
LICO3 Listera cordata
LOCI Lonicera ciliosa
LOCR Lotus crassifolius
LOMI Lotus micranthus
LOPU Lotus purshianus
LULA Lupinus latifolius
MAGR Madia gracilis
MOPE Montia perfoliata
MOSI Montia sibirica
MYDI Myosotis discolor
NEPA Nemophila parviflora
OSCH Osmorhiza chilensis
PEFR2 Petasites frigidus
PILA Pinus lambertiana
POLYG Polygonum
PODO Polygonum douglasii
POMU Polystichum munitum
PRVU Prunella vulgaris
PREM Prunus emarginata
PSME Pseudotsuga menziesii
PTAQ Pteridium aquilinum
PYAS Pyrola asarifolia
PYPI Pyrola picta
RAUN2 Ranunculus uncinatus
RHMA Rhododendron macrophyllum
RIBES Ribes
RILO Ribes lobbii
ROGY Rosa gymnocarpa
RUBUS Rubus
RULE Rubus leucodermis
RUNI Rubus nivalis
RUPA Rubus parviflorus
RUUR Rubus ursinus
SACE Sambucus cerulea
SEJA Senecio jacobaea
SESY Senecio sylvaticus
SIAN2 Silene antirrhina
SOAR Sonchus arvensis
STELL Stellaria
STCA2 Stellaria calycantha
TABR Taxus brevifolia
THPL Thuja plicata
TITR Tiarella trifoliata
TRLA2 Trientalis latifolia
TROV Trillium ovatum
TRCA Trisetum canescens
TRCE Trisetum cernuum
TSHE Tsuga heterophylla
VAME Vaccinium membranaceum
VAPA Vaccinium parvifolium
VAHE Vancouveria hexandra
VEAR Veronica arvensis
VESE Veronica serpyllifolia
VISE Viola sempervirens
WHMO Whipplea modesta
CACH Castanopsis chrysophylla
EBAU Eburophyton austiniae
SADO Satureja douglasii
SMST Smilacina stellata
EPAN Epilobium angustifolium
SYMO Symphoricarpos mollis
GNMI Gnaphalium microcephalum
LUCA2 Luzula campestris
UNKN unknown plant
HERB Herb
TREE Tree
SHRUB Shrub
MOSS Moss
LICHEN Lichen
GRASS Grass
LITTER Litter
CWD Coarse woody debris
BARE Bare ground
LOG Log
STONE Stone
BUTT Tree butt
STUMP Tree stump
SNAG Snag
UNKN2 unknown plant 2
ROOT Root

Enumerated domain for attribute: STRATUM

Codes for STRATUM
Code Description
O Overstory layer
S Subcanopy layer
U Understory layer
N Not applicable; STRATUM applies only to tree species in the pre-harvest sample (1990)

Enumerated domain for attribute: ALTEQU

Codes for ALTEQU
Code Description
1 primary equation
2 second alternative equation
3 third alternative equation
4 fourth alternative equation

Enumerated domain for attribute: BURN_CLASS

Codes for BURN_CLASS
Code Description
B Burned
PB Partially burned
UB Unburned