Grier, C. C.; Cole, D. W.; Dyrness, C. T.; Fredriksen, R. L. 1974. Nutrient cycling in 37- and 450-year-old Douglas-fir ecosystems. In: Waring, R. H.; Edmonds, R. L., eds. Integrated research in the coniferous forest biome. Bulletin 5. Seattle, WA: Coniferous Forest Biome: 21-34.
Biomass and nitrogen, phosphorus, potassium, and calcium distribution,and biogeochemical and stand nitrogen, phosphorus, potassium, and calciumbudgets were determined for 37- and 450-year-old Pseudotsuga menziesii
(Mirb.) Franco stands in the U.S. Pacific Northwest. Biomass of the
450-year-old stand is greater, but annual growth is less than that of
the 37-year-old stand. About 50% of the annual growth and over 50% of
the nutrient uptake and return in the 450-year-old stand occurs in sub-
ordinate vegetation compared with less than 15% in the 37-year-old stand.Chemical differences in soil parent material between the two stands are
reflected in both the biogeochemical and stand nutrient cycles.